Carex of Arkansas

Revised September 24, 2003

Carex of Arkansas: an annotated list

This page provides a list of all Arkansas Carex sedges. On March 9, 2003, I started detailed work on the Section Laxiflorae. With that beginning, I plan to treat each group of Carex sedges by the section to which they belong. Most info here comes from my draft manuscript on the genus in Arkansas; that document runs about 115 pages, and contains much info not available here. It will take considerable time to make it available on line. But it has to be done someday! If you want a copy of that manuscript, please email me.

Taxonomic problems have settled out within Carex, as I've worked on the genus in Arkansas since 1991. Yet, within the last two years (2001-2002), additional species new-to-science have been described which occur in Arkansas. Further work on southeastern Carex sedges is expected to turn up more "splinter" species. So far, as I've studied them, these new species seem to prove valid, based on differences in habitat, morphology, and the fact that even a non-university based botanist-specialist like myself can usually recognize them with ease.

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Based on Hyatt (1998) and Smith (1994). Will be revised after further research. Starting with the Laxiflorae in March 2003, I plan to move details on each species into a separate page for each section. In the cases of small sections with few species, more than one section may be treated on a single page.

Carex abscondita Mackenzie

...This sedge occurs much more frequently than collections indicate. Find it by looking along medium to larger sized streams along natural levees. It frequents nearby valleys also but is easily overlooked as the perigynia are hidden in the leaves.

Carex aggregata Mackenzie

Carex alata Torrey

Carex albicans Willdenow var. albicans

...Common in the Ozarks and Ouachitas, this sedge comes up early and frequents bluffs, rock outcrops, ridgetops, and slopes. Its told from most sedges by the fine hairs on the perigynia, which you can see with a hand lens. Its densely cespitose, but sometimes has short rhizomes less than a few centimeters long.

Carex albicans Willdenow var. australis (L. H. Bailey) Rettig

...This plant occurs more commonly in the south, but is reportedly "rare" throughout its range, the southeast United States. It ain't so! Its just infrequent because its infrequently collected, and blooms earlier than most sedges. Its similar to the var. albicans, but has long rhizomes, usually.

Carex albolutescens Schweinitz

Carex albursina Sheldon of section Laxiflorae

Carex amphibola Steudel

Carex annectens Bicknell var. annectens

Carex arkansana L. H. Bailey

Carex atlantica L. H. Bailey spp. atlantica

Carex atlantica L. H. Bailey spp. capillacea (L. H. Bailey) Reznicek

Carex austrina (Small) Mackenzie

Carex basiantha Schkuhr

Carex bicknellii Britton

Known as the "Texas form" something's out there underCarex bicknellii Britton that's not that species exactly.

Carex bicknellii Britton var. opaca F. J. Hermann

Carex blanda Dewey of section Laxiflorae

Carex brevior (Dewey) Mackenzie ex Lunell

Carex buffaloensis spp. nov.

This Carex is another undescribed entity.

Carex bromoides Schkuhr var. bromoides

Carex bulbostylis Mackenzie

Carex bullata Schkuhr

Carex bushii Mackenzie

Carex careyana Torrey

Carex caroliniana Schweinitz

Carex cephalophora Muhlenberg ex Willdenow

Carex cherokeensis Schweinitz

Carex communis L. H. Bailey

Carex comosa Boott

Carex complanata Torrey & Hooker

Carex conjuncta Boott

Carex corrugata Fernald

Carex crawei Dewey

Carex crebriflora Wiegand of section Laxiflorae

Carex crinata Lamarck

Carex crus-corvi Shuttleworth ex Kuntze

Carex davisii Schweinitz & Torrey

Carex debilis Michaux var. debilis

Carex debilis Michaux var. pubera Gray

Carex decomposita Muhlenberg of section Heleoglochin

Carex digitalis Willdenow var. asymetrica Fernald

Carex digitalis Willdenow var. digitalis

Carex digitalis Willdenow var. macropoda Fernald

Carex eburnea Boott

Carex emoryi Dewey

Carex festucacea Schkuhr

Carex fissa Mackenzie var. fissa

Carex flaccosperma Dewey

Carex frankii Kunth

Carex gigantea Rudge

Carex glaucescens Elliott

Carex glaucodea Tuckerman in Olney in A. Gray

Carex gracilescens Steudel of section Laxiflorae

Carex gracillima Schweinitz

Carex granularis Muhlenberg

Carex gravida L. H. Bailey

Carex grayi Carey

Carex grisea Wahlenberg

Carex hirsutella Mackenzie

Carex hirtifolia Mackenzie

Carex hitchcockiana Dewey

Carex hyalina Boott

Carex hyalinolepis Steudel (Shoreline Sedge)

[C. impressa (S. H. Wright) Mack.] OBL. Large sterile colonies of this plant often exist, but just as often the culms hide among the much longer leaves, protected from the would be collector by cottonmouth snakes and several inches of water, leaving the collector on the shore. In addition, large long rhizomes make hands in muck the fastest way of retriving a complete collection; in reality collectors easily collect rhizomes in wet weather by loosing the soil around them. Yanked culms come up rootless and rhizomeless. The name Shoreline Sedge fits this species well. This plant dominates a narrow strip along roadside ditches; natural habitats include wet woods and in prairies as indicated on Sundell 6226 at UAM: "dominated monotypic patches about 15-25 feet in diameter, in wet areas of Warren Prairie." To the inexperienced, the plant looks similar to Carex cherokeensis, which has much shorter, very stout rhizomes with many culms which form circular colonies in mesic but much drier sites than C. hyalinolepis. Once learned, these two taxa are almost unmistakeable with mature collections, especially with the rhizomes present in the collection. It is scattered in the southeastern half of Arkansas, but should be looked for in the Arkansas river valley and in unsearched counties within the existing range, while it is unexpected in the Ouachitas and Ozarks generally. "Ranges from Florida to Texas, north to New Jersey and Pennsylvania, Ontario, Ohio, Michigan, Wisconsin, Iowa, and Nebraska" (Steyermark 1963).

Carex hystericina Muhlenberg ex Willdenow

Carex interior L. H. Bailey

Carex intumescens Rudge

Carex jamesii Schweinitz

Carex joorii L. H. Bailey

Carex kraliana Naczi & Bryson of section Laxiflorae

Carex laevivaginata (Kukenth) Mackenzie

Carex latebracteata Waterfall

Carex laxiculmis Schweinitz var. copulata (Bailey) Mackenzie

Carex laxiculmis Schweinitz var. laxiculmis

Carex laxiflora Lamarck of section Laxiflorae

Carex leavenworthii Dewey

Carex leptalea Wahlenberg var. harperi

Carex leptalea Wahlenberg var. leptalea

Carex longii Mackenzie

Carex louisianica L. H. Bailey

Carex lupuliformis Sartwell

Carex lupulina Muhlenberg

Carex lurida Wahlenberg

Carex meadii Dewey

Carex mesochorea Muhlenberg ex Willdenow

Carex microdonta Torrey & Hooker

Carex molesta Mackenzie

Carex molestiformis Reznicek & P. Rothrock

Carex muhlenbergii Willdenow var. enervis Boott

Carex muhlenbergii Willdenow var. muhlenbergii

Carex muskingumensis Schweinitz

Carex nigromarginata Schweinitz

Carex normalis Mackenzie

Carex oklahomensis Mackenzie

Carex oligocarpa Schkuhr

Carex ouachitana Kral, Manhart, & Bryson

Carex oxylepis Torrey & Hooker var. oxylepis

Carex oxylepis Torrey & Hooker var. pubescens Underwood

Carex ozarkana P. Rothrock & Reznicek

Carex pellita Willdenow (Wooly Sedge)

[Carex lanuginosa Michaux] OBL. Lawrence County provides the only known Arkansas location of this plant (Russell McNalty 321 STAR, ! peh, ! E. B. Smith, ! Barney Lipscomb; collected roadside 2 miles east of Walnut Ridge on Hwy. 25 in water on April 16, 1968). Demaree (1943) listed it for Arkansas, but no voucher of his report has been located yet. A Missouri site occurs in a large sedge and grass dominated area of a huge area of sinks a few miles from Arkansas. It could occur in north central Arkansas within similar habitat. The author continues to seek out areas with sinkhole ponds, to search for this and other species. Most sinks found to date tend to be dry holes rather than sunny wet ponds. "Ranges from Quebec to British Columbia south to Virginia, Tennessee, Arkansas, Oklahoma, Texas, New Mexico, Arizona, and California" (Steyermark 1963). The single Arkansas voucher has a 20 cm long rhizome with nodes less than 1 cm apart, rooting at the nodes with 8 cm long fibrous roots. Most leaves are missing, cut off about 3 cm tall, so leaf characters are lost. The 26 cm of culm contains raised nerves, and a 26 cm long bract below the spikelets, about 3-4 mm wide toward the base and a less than 5 mm long sheath. The ligule is not visible without tearing the voucher. The specimen has a terminal staminate spikelet, 2 subterminal (either immature or aborted) pistillate spikelets, and two lower, mature pistillate spikelets. The 3.5 cm long staminate spikelet has acute scales, with those at the tip and base having awns up to 0.3-0.4 mm long. Tinges of a red wine color remain in this spikelet. At its base is a 1.3 cm long pistillate spikelet, and 1.4 cm lower another 1.3 cm long aborted pistillate spikelet. About 5 cm lower on the culm is the mature pistillate spikelet and the second, about 6 cm below it. These are 2.6 and 3.7 cm long, respectively. The pistillate scales have scabrous awns 1.5-3.0 mm long, the base of the awn continuing centrally down the scale as a green nerve to 0.4 mm wide flanked by a thin, translucent wing, 0.5-0.7 mm wide and tinged brown (purple in life?) outer edge. A fine, rough hairness covers the perigynia. The glabrous 0.5-0.8 mm long divisions of the beaks of the perigynia sit atop the 0.8-1.0 mm long undivided portion of the beaks. The teardrop shaped perigynia run about 4 mm long, including the beak. I estimated 65 perigynia in a single spikelet. This collection contains a second immature culm of perhaps C. meadii or C. blanda.

Carex pensylvanica Lamarck

Carex planispicata Naczi

Carex planostachys Kunze

Carex prasina Wahlenberg

Carex radiata (Wahlenberg) Small

Carex reniformis (L. H. Bailey) Small

Carex retroflexa Willdenow

Carex rosea Schkuhr

Carex scoparia Schkuhr ex Willdenow

Carex seorsa Howe

Carex shortiana Dewey

Carex socialis Mohlenbrock & Schwegmann

Carex sparganioides Muhlenberg

Carex squarrosa Linnaeus

Carex striatula Michaux of section Laxiflorae

Carex styloflexa of section Laxiflorae

Carex stricta Lamarck

Carex suberecta (Olney) Britton

Carex swanii (Fernald) Mackenzie

Carex texensis L. H. Bailey

Carex torta Boott

Carex triangularis Boeckler

Carex tribuloides Wahlenberg

Carex typhina Michaux

Carex umbellata Schkuhr

Carex virescens Muhlenberg

Carex vulpinoidea Michaux

Carex willdenowii Willdenow var. willdenowii

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References

Hyatt, Philip E. 1999. Arkansas Carex (Cyperaceae): a preliminary list [or some such title]. Sida. Correct citation to be added later.

Smith, Edwin B. 1994. Keys to the Flora of Arkansas. The University of Arkansas Press. Fayetteville. 363 pages.

Yatskievych, George. 1999. [revised Missouri flora volume 1]


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